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Colonel Lovok was a member of the Tal Shiar. He worked along side with the Obsidian Order in their plan to destroy the Founders.
At some point Lovok was replaced by a Changeling, unbeknownst to his colleagues and conspirators. His presence in the plot allowed the Dominion to have enough forewarning to evacuate the Founders' homeworld and set a trap for the joint Tal Shiar-Obsidian Order fleet.
The Changeling revealed itself to Odo and Garak when it assisted them in escaping from his warbird during the battle. (DS9: "The Die is Cast")